Team’s described species
|
Described taxa |
Publication |
Described from |
Current accepted name (if changed) |
|
Lewinskya transcaucasica Eckstein, Garilleti & F.Lara |
(Eckstein et al. 2018) |
Caucasus |
|
|
L. lamyana F.Lara, Garilleti, Draper & Mazimpaka |
(Lara et al. 2018) |
Mediterranean |
|
|
L. scissa Vigalondo, F.Lara & Garilleti |
(Vigalondo et al. 2020) |
Canary Is. |
|
|
L. arida Vigalondo, F.Lara & Garilleti |
(Vigalondo et al. 2020) |
W North America |
|
|
L. pacifica Vigalondo, F.Lara & Garilleti |
(Vigalondo et al. 2020) |
W North America |
|
|
L. pseudoaffinis Vigalondo, F.Lara & Garilleti |
(Vigalondo et al. 2020) |
W North America |
|
|
Orthotrichum ibericum F.Lara & Mazimpaka |
(Lara & Mazimpaka 1993) |
Mediterranean |
Lewinskya iberica (F.Lara & Mazimpaka) F.Lara, Garilleti & Goffinet |
|
O. macrocephalum F.Lara, Garilleti & Mazimpaka |
(Lara et al. 1994) |
Mediterranean |
|
|
O. tortidontium F.Lara, Garilleti & Mazimpaka |
(Lara et al. 1996) |
Mediterranean |
Lewinskya tortidontia (F.Lara, Garilleti & Mazimpaka) F.Lara, Garilleti & Goffinet |
|
O. lewinskyae F.Lara, Garilleti & Mazimpaka |
(Garilleti et al. 1997) |
Mediterranean |
Orthotrichum scanicum Grönvall |
|
O. vittii F.Lara, Garilleti & Mazimpaka |
(Lara et al. 1999) |
Mediterranean |
|
|
O. casasianum F.Lara, Garilleti & Mazimpaka |
(Mazimpaka et al. 1999) |
Mediterranean |
|
|
O. handiense F.Lara, Garilleti & Mazimpaka |
(Lara et al. 1999) |
Canary Is. |
|
|
O. hispanicum F.Lara, Garilleti & Mazimpaka |
(Lara et al. 2000b) |
Mediterranean |
|
|
O. underwoodii F.Lara, Garilleti & Mazimpaka |
(Garilleti et al. 2001) |
W North America |
|
|
O. speciosum var. brevisetum F.Lara, Garilleti & Mazimpaka |
(Draper et al. 2003) |
Mediterranean |
Lewinskya breviseta (F.Lara, Garilleti & Mazimpaka) F.Lara, Garilleti & Goffinet |
|
O. anodon F.Lara, Garilleti & Mazimpaka |
(Garilleti et al. 2006) |
W North America |
|
|
O. galiciae F.Lara, Garilleti & Mazimpaka |
(Lara et al. 2006) |
E Africa |
Lewinskya galiciae (F.Lara, Garilleti & Mazimpaka) F.Lara, Garilleti & Goffinet |
|
O. spiculatum F.Lara, Garilleti & Mazimpaka |
(Lara et al. 2007) |
C Sudamérica |
|
|
O. norrisii F.Lara, Medina & Garilleti |
(Medina et al. 2008) |
W North America |
|
|
O. karoo F.Lara, Garilleti & Mazimpaka |
(Lara et al. 2009) |
S Africa |
|
|
O. mazimpakanum Garilleti & F.Lara |
(Garilleti et al. 2011) |
W North America |
|
|
O. pilosissimum Medina, F.Lara & Garilleti |
(Medina et al. 2011) |
W North America |
|
|
O. cambrense Bosanquet & F.Lara |
(Bosanquet & Lara 2012) |
Europa |
|
|
O. confusum Medina, F.Lara & Garilleti |
(Medina et al. 2012) |
W North America |
|
|
O. persimile F.Lara, Medina, Garilleti & Mazimpaka |
(Medina et al. 2012) |
W North America |
|
|
O. comosum F.Lara, Medina & Garilleti |
(Medina et al. 2013) |
Mediterranean |
|
|
O. cucullatum F.Lara, Medina & Garilleti |
(Medina et al. 2013) |
W North America |
|
|
O. franciscanum F.Lara, Medina & Garilleti |
(Medina et al. 2013) |
W North America |
|
|
Ulota gigantospora F.Lara, Caparrós & Garilleti |
(Caparrós et al. 2011) |
E Asia |
|
|
U. yunnanensis F.Lara, Caparrós & Garilleti |
(Caparrós et al. 2011) |
E Asia |
|
|
U. billbuckii Garilleti, Mazimpaka & F.Lara |
(Garilleti et al. 2012) |
Patagonia |
|
|
U. streptodon Garilleti, Mazimpaka & F.Lara |
(Garilleti et al. 2012) |
Patagonia |
|
|
U. larrainii Garilleti, Mazimpaka & F.Lara |
(Garilleti et al. 2015) |
Patagonia |
|
|
U. maltana Garilleti & F.Lara |
(Garilleti et al. 2020) |
Patagonia |
|
|
Zygodon catarinoi C.Garcia, F.Lara, Sèrgio & Sim-Sim |
(Garcia et al. 2006) |
Mediterranean |
|
|
Z. chilensis Calabrese & F.Lara |
(Calabrese et al. 2006) |
Patagonia |
Australoria chilensis (Calabrese & F.Lara) F.Lara, Garilleti & Draper |
|
Species resurrected from synonymy |
Publication |
|
Lewinskya shawii (Wilson) F. Lara, Garilleti & Goffinet |
Mazimpaka et al. 2000 |
|
Lewinskya fastigiata (Bruch ex Brid.) Vigalondo, F. Lara & Garilleti |
Vigalondo et al. 2019 |
|
Lewinskya leptocarpa (Bruch & Schimp. ex Müll. Hal.) Vigalondo, F. Lara & Garilleti |
Vigalondo et al. 2019 |
|
Orthotrichum columbicum Mitt. |
Medina et al. 2012 |
|
Orthotrichum cylindrocarpum Lesq. |
Medina et al. 2013 |
|
Pulvigera howei (Renauld & Cardot) F. Lara, Draper & Garilleti |
Lara et al. 2020 |
|
Pulvigera papillosa (Hampe) F. Lara, Draper & Garilleti |
Lara et al. 2020 |
|
Pulvigera pringlei (Müll. Hal.) F. Lara, Draper & Garilleti |
Lara et al. 2020 |
|
Ulota macrodontia Dusén ex Malta |
Garilleti et al. 2016 |
|
Ulota crispula Bruch |
Caparrós et al. 2016 |
|
Ulota intermedia Schimp. |
Caparrós et al. 2016 |
|
Ulota pygmaeothecia (Müll.Hal.) Kindb. |
Muñoz-Puelles et al. 2017 |
|
Ulota glabella Mitt. |
Garilleti et al. 2020 |
|
Supraspecific taxa proposed |
Publication |
|
Tribe |
|
|
Codonoblepharonteae F. Lara, Garilleti & Draper |
Aguado-Ramsay et al. 2022 |
|
Subtribes |
|
|
Lewinskyinae F.Lara, Garilleti & Draper |
Draper et al. 2021 |
|
Orthotrichinae F.Lara, Garilleti & Draper |
Draper et al. 2021 |
|
Genera |
|
|
Atlantichella F.Lara, Garilleti & Draper |
Draper et al. 2021 |
|
Australoria F.Lara, Garilleti & Draper |
Draper et al. 2021 |
|
Lewinskya F.Lara, Garilleti & Goffinet |
Lara et al. 2016 |
|
Rehubryum F.Lara, Garilleti & Draper |
Draper et al. 2022 |
List of used taxonomical characters
Mediterranean Orthotrichaceae
The Mediterranean basin is home to a high diversity of Orthotrichaceae (Ros et al., 2013, Hodgetts & Lockhart 2020). In this biodiversity hotspot (CEPF), 68 species and varieties of this family are found, 83 % of all those known in Europe, North Africa and the Near East. Of these, 15 species are endemic or almost exclusively found in this biogeographic area.
|
Europaean taxa |
Mediterranean |
Europaean taxa |
Mediterranean |
|
Atlantichella calvescens |
✓ |
O. microcarpum |
(✓) |
|
Codonoblepharon forsteri |
✓ |
O. moravicum |
|
|
Lewinskya acuminata |
✓ |
O. pallens |
✓ |
|
L. affinis |
✓ |
O. patens |
(✓) |
|
L. breviseta |
✓ |
O. pellucidum |
|
|
L. elegans |
|
O. philibertii |
✓ |
|
L. fastigiata |
✓ |
O. pulchellum |
✓ |
|
L. iberica |
✓ |
O. pumilum |
✓ |
|
L. iwatsukii |
|
O. rivulare |
✓ |
|
L. laevigata |
✓ |
O. rogeri |
(✓) |
|
L. lamyana |
✓ |
O. scanicum |
✓ |
|
L. pylaisii |
|
O. schimperi |
✓ |
|
L. rupestris |
✓ |
O. shevockii |
✓ |
|
L. shawii |
✓ |
O. sibiricum |
|
|
L. sordida |
|
O. sprucei |
✓ |
|
L. speciosa |
✓ |
O. stellatum |
(✓) |
|
L. striata |
✓ |
O. stramineum |
✓ |
|
L. tortidontia |
✓ |
O. tenellum |
✓ |
|
L. transcaucasica |
|
O. urnigerum |
(✓) |
|
L. vladikavkana |
(✓) |
O. vittii |
✓ |
|
Nyholmiella gymnostoma |
✓ |
Plenogemma phyllantha |
(✓) |
|
N. obtusifolia |
✓ |
Pulvigera lyellii |
✓ |
|
Orthotrichum alpestre |
✓ |
Ulota bruchii |
✓ |
|
O. anomalum |
✓ |
U. coarctata |
(✓) |
|
O. bistratosum |
✓ |
U. crispa |
✓ |
|
O. callistomum |
|
U. crispula |
✓ |
|
O. cambrense |
|
U. curvifolia |
|
|
O. casasianum |
✓ |
U. drummondii |
(✓) |
|
O. columbicum |
✓ |
U. hutchinsiae |
✓ |
|
O. comosum |
✓ |
U. intermedia |
✓ |
|
O. consobrinum |
✓ |
U. macrospora |
(✓) |
|
O. crenulatum |
✓ |
U. rehmannii |
(✓) |
|
O. cupulatum var. cupulatum |
✓ |
Zygodon catarinoi |
✓ |
|
O. cupulatum var. fuscum |
(✓) |
Z. conoideus var. conoideus |
✓ |
|
O. cupulatum var. riparium |
✓ |
Z. conoideus var. lingulatus |
|
|
O. dagestanicum |
|
Z. dentatus |
(✓) |
|
O. dentatum |
(✓) |
Z. gracilis |
(✓) |
|
O. diaphanum |
✓ |
Z. rupestris |
✓ |
|
O. handiense |
✓ |
Z. sibiricus |
|
|
O. hispanicum |
✓ |
Z. stirtonii |
(✓) |
|
O. macrocephalum |
✓ |
Z. viridissimus |
✓ |
Keys for Mediterranean Orthotrichaceae:
Orthotrichaceae in Handbook of mosses the Iberian Peninsula and the Balearic Islands (Updated Brugués & Ruiz 2020)
- Key for Ulota s.l. (Muñoz-Puelles et al. 2018)
- Orthotrichaceae in Guerra J, Cano MJ & Brugués, M (eds), Flora Briofítica Ibérica, vol V. (AVAILABLE AT: Lara et al. 2014)
- Key to Orthotrichum s.l. in the Mediterranean area (Lara et al. 2009)
- Key for Zygodon s.l. (Calabrese & Muñoz 2008)
Basic bibliography on Orthotrichaceae
Allen B. 2002. Moss flora of Central America. Part 2. Encalyptaceae-Orthotrichaceae. Monographs in Systematic Botany from the Missouri Botanical Garden 90: 1–699.
Calabrese GM. 2006. A taxonomic revision of Zygodon (Orthotrichaceae) in southern South America. The Bryologist 109: 453–509.
Draper I, Garilleti R, Calleja JA, Flagmeier M, Mazimpaka V, Vigalondo B & Lara F. 2021. Insights into the evolutionary history of the subfamily Orthotrichoideae (Orthotrichaceae, Bryophyta): new and former supra-specific taxa so far obscured by prevailing homoplasy. Frontiers in Plant Sciences 12: 629035.
Draper, I., T. Villaverde, R. Garilleti, G. Burleigh, S. McDaniel, V. Mazimpaka, J.A Calleja, & F. Lara. 2022. An NGS-based phylogeny of Orthotricheae (Orthotrichaceae, Bryophyta) with the proposal of the new genus Rehubryum from Zealandia. Frontiers in Plant Science 13: 882960.
Goffinet B, Shaw AJ, Cox CJ, Wickett NJ & Boles SB. 2004. Phylogenetic inferences in the Orthotrichoideae (Orthotrichaceae, Bryophyta) based on variation in four loci from all genomes. Monographs in Systematic Botany from the Missouri Botanical Garden 98: 270–289.
Goffinet B & Vitt DH. 1998. Revised generic classification of the Orthotrichaceae based on a molecular phylogeny and comparative morphology. In: Bates JW, Ashton NW, Duckett JG, eds. Bryology for the Twenty-first Century. Leeds: Maney Publishing and the British Bryological Society, 143–160.
Lara F, Draper I, Flagmeier M, Calleja JA, Mazimpaka V & Garilleti R. 2020. Let ’s make Pulvigera great again: re-circumscription of a misunderstood group of Orthotrichaceae that diversified in North America. Botanical Journal of the Linnean Society 193: 180–206.
Lara F, Garilleti R, Goffinet B, Draper I, Medina R, Vigalondo B & Mazimpaka V. 2016. Lewinskya, a new genus to accommodate the phaneroporous and monoicous taxa of Orthotrichum (Bryophyta, Orthotrichaceae). Cryptogamie, Bryologie 37: 361–382.
Lara F, Garilleti R, Mazimpaka V & Guerra J (Eds.). 2014. Orthotrichaceae. In: J. Guerra, M. J. Cano & M. Brugués (eds.) Flora Briofítica Ibérica, vol. V. UMU / SEB. Murcia. Pp.: 15-135.
Lewinsky J. 1993. A synopsis of the genus Orthotrichum Hedw. (Musci: Orthotrichaceae). Bryobrothera 2: 1–59.
Lewinsky-Haapasaari J. 1995. Illustrierter bestimmungsschlüssel zu den europäischen Orthotrichum-Arten. Meylania: 3–56.
Malta N. 1926. Die Gattung Zygodon Hook. et Tayl. Ein monographischen Studie. Riga: Latvijas Universitates.
Malta N. 1927. Die Ulota-Arten Süd-Amerikas. Acta Horti Botanici Universitatis Latviensis 2: 165–208.
Malta N. 1933. A survey of the Australasian species of Ulota. Acta Horti Botanici Universitatis Latviensis 7: 1–24.
Plášek V, Sawicki J, Ochyra R, Szczecińska M & Kulik T. 2015. New taxonomical arrangement of the traditionally conceived genera Orthotrichum and Ulota (Orthotrichaceae, Bryophyta). Acta Musei Silesiae, Scientiae Naturales 64: 169–174.
Vitt DH. 1973. a revision of the genus Orthotrichum in North America, north of Mexico. Bryophytorum Bibliotheca 1: 1–165.
OTHER INTERESTING REFERENCES
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Atwood JJ. 2009. A taxonomic revision of Schlotheimia subg. Stegotheca (Orthotrichaceae). The Bryologist 112: 290–300.
Blockeel TL. 2017. The Ulota crispa group in Britain and Ireland, with notes on other species of the genus. Field Bryology 117: 8–19.
Bosanquet SDS. 2009. Orthotrichum – Britain’s bristle-mosses. British Wildlife 20: 187–194.
Calabrese GM & Muñoz J. 2008. Zygodon (Orthotrichaceae) in the Iberian Peninsula. The Bryologist 111: 231–247.
Caparrós R, Lara F, Draper I, Mazimpaka V & Garilleti R. 2016. Integrative taxonomy sheds light on an old problem: The Ulota crispa complex (Orthotrichaceae, Musci). Botanical Journal of the Linnean Society 180: 427–451.
Crum HA & Anderson LE. 1981. Mosses of Eastern North America, Vol. 1.
Eddy A. 1996. Orthotrichaceae. In: A handbook of Malesian mosses. Volume 3. Splachnobryaceae to Leptosmataceae. London: HMSO, 12–96.
Fedosov VE. 2018. The family Orthotrichaceae (Bryophyta) in flora of Russia: results of revision and biogeographical review (Семейство Orthotrichaceae (Bryophyta) во флоре России: итоги ревизии и биогеографический обзор). Novosti sistematiki nizshikh rastenii 52: 519–534.
Fedosov VE & Ignatova EA. 2018. On the genus Ulota (Orthotrichaceae, Bryophyta) in Russia. Novosti sistematiki nizshikh rastenii 52: 141–171.
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Goffinet B, Bayer RJ & Vitt DH. 1998. Circumscription and phylogeny of the Orthotrichales (Bryopsida) inferred from rbcL sequence analyses. American Journal of Botany 85: 1324–1337.
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Lewinsky J. 1987. Orthotrichum (Orthotrichaceae) in South America 2. Taxonomic revision of taxa with superficial stomata. Memoirs of the New York Botanical Garden 45: 326–370.
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